Cryptantha Sections & Series

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Adelinia Amsinckia Amsinckiopsis Andersonglossum Cryptantha Dasynotus Eremocarya Greeneocharis Gruvelia Harpagonella Johnstonella Oncaglossum Oreocarya Pectocarya Plagiobothrys Simpsonanthus Sonnea

Systematics of Amsinckiinae (Boraginaceae): The Popcorn Flowers

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Cryptantha Lehmann ex G.Don

Gen. Hist. 4(1): 373 (1837)

Pronunciation/Pronunciación:
Cryp-tán-tha
Etymology/Etimología:

krypto/crypto, "hidden / econdido" + anthos, "flower / flor," after the cleistogamous flowers in the type species / de las flores cleistógamas en la especies tipo
Synonyms/Sinónimos:












Echinospermum Sw. ex Lehm.
Eremocarya Greene
Eritrichium Schrad. ex Gaudin
Greeneocharis Gürke & Harms
Hemisphaerocarya Brand
Johnstonella Brand
Krynitzkia Fisch. & C.A. Meyer, Index Seminum [St. Petersburg] 7: 52. 1841
Lithospermum L.
Myosotis L.
Oreocarya Greene
Piptocalyx Torr., ined.
Rochelia Rch.
Wheelerella G.B. Grant, ined.]

Type Species/EspeciesTipo:

Cryptantha glomerata Lehmann ex G.Don, Gen. Hist. 4(1): 373 (1837)
      Lectotype chosen by Johnston 1935, J. Arnold Arbor. 16(2): 196. 1935 [24 Apr 1935]

Distribution of Cryptantha:
cryptantha distribution 1
cryptantha distribution 2
LEFT: GBIF 2019: https://www.gbif.org. Accessed 24 Dec 2019.
RIGHT: POWO. 2019. Plants of the World Online. http://www.plantsoftheworldonline.org. Green=native. Accessed 24 Dec 2019.




TOTAL NUMBER OF TAXA IN GENUS:
109
species and 124 minimally ranked taxa
FULL SYNONOMY HERE

NORTH AMERICAN TAXA
[SA = also in South America]

63 species (77 taxa) of Cryptantha in North America
63 especies (77 taxones) de Cryptantha de Norteamericana
Johnston's 1925 Treatment


Listing by Series [*=Series placement not recognized by Johnston]
Back to Alphabetical Listing

SOUTH AMERICAN TAXA
[NA = also in North America]

47 species (48 taxa) of Cryptantha in South America
47 especies (48 taxones) de Cryptantha de Sudamericana
Johnston's 1927 Treatment
Johnston's 1929 Treatment

Listing by Series [*=Series placement not recognized by Johnston]
Back to Alphabetical Listing

TAXON/TAXÓN                                                                 COUNTRY/PAÍS
Series Affines (Johnston 1925)    
Cryptantha affinis (A.Gray) Greene
Canada USA
Cryptantha clevelandii Greene *
Mexico USA
   var. clevelandii
Mexico USA
   var. florosa
Mexico USA
Cryptantha corollata (I.M.Johnston) I.M.Johnston *
  USA
Cryptantha decipiens (M.E.Jones) A.Heller *
Mexico USA
Cryptantha glomeriflora Greene 4.3 *
USA
Cryptantha hispidula Greene ex Brand  SE *
USA
Cryptantha microstachys (Greene ex A.Gray) Greene * 
Mexico USA
Cryptantha rattanii Greene 4.3 *
USA
     
Series Ambiguae (Johnston 1925)    
Cryptantha ambigua (A.Gray) Greene Canada USA
Cryptantha crinita Greene 1B.2
USA
Cryptantha mariposae I. M. Johnston 1B.3  SE
  USA
Cryptantha torreyana (A.Gray) Greene
Canada USA
   var. pumila (A. Heller) I.M.Johnston
USA
   var. torreyana
Canada USA
Cryptantha watsonii (A.Gray) Greene *
Canada USA
     
Series Barbigerae (Johnston 1925)    
Cryptantha arenophila Rebman & M.G.Simpson
Mexico  
Cryptantha barbigera (A.Gray) Greene
Mexico USA
   var. barbigera
Mexico USA
   var. fergusoniae J.F.Macbride
USA
Cryptantha catalinensis M.G.Simpson & Rebman
USA
Cryptantha clementina Rebman & M.G.Simpson
USA
Cryptantha ganderi I.M.Johnston 1B.1  *
  Mexico USA
Cryptantha grandiflora Rydberg
USA
Cryptantha hendersonii (A.Nelson) Piper ex J.C.Nelson
USA
Cryptantha intermedia (A.Gray) Greene  
Canada? Mexico USA
  var. johnstonii J.F.Macbride
Mexico USA
  var. intermedia  WI/IN 
Mexico USA
Cryptantha kinkiensis Rebman & M.G.Simpson
USA
Cryptantha milobakeri I.M.Johnston  SI
USA
Cryptantha patula Greene *
Mexico  
Cryptantha wigginsii I.M.Johnston 1B.2
Mexico USA
     
Series Dumertoreae *     
Cryptantha dumetorum (Greene ex A.Gray) Greene
USA
     
Series Echinellae *    
Cryptantha echinella Greene
USA
Cryptantha incana Greene 1B.3
  USA
     
Series Flaccidae (Johnston 1925)  
Cryptantha flaccida (Douglas ex Lehmann) Greene  WI
USA
Cryptantha rostellata (Greene) Greene 4.2
USA
Cryptantha simulans Greene *
USA
Cryptantha sparsiflora (Greene) Greene
USA
Cryptantha spithamaea I.M.Johnston 1B.3
USA
     
Series Foliosae *    
Cryptantha foliosa (Greene) Greene
Mexico  
Cryptantha traskiae I.M.Johnston 1B.1
USA
     
Series Leiocarpae (Johnston 1925)    
Cryptantha dissita I.M.Johnston 1B.2  BE/SI
USA
Cryptantha hispidissima Greene
USA
Cryptantha juniperensis R.B.Kelley & M.G.Simpson *
  Mexico USA
Cryptantha leiocarpa (Fischer & C.A.Meyer) Greene
  USA
Cryptantha nemaclada Greene
USA
     
Series Micromereae *     
Cryptantha micromeres (A.Gray) Greene
Mexico USA
     
Series Pterocaryae (Johnston 1925)    
Cryptantha cycloptera (Greene) Greene 
  Mexico USA
   f. truncata M.E.Mabry & Rebman
  USA
Cryptantha gracilis Osterhout *
USA
Cryptantha mohavensis (Greene) Greene *
USA
Cryptantha oxygona (A.Gray) Greene
USA
Cryptantha pterocarya (Torrey) Greene 
Mexico USA
  var. pterocarya
Mexico USA
     f. pseudocycloptera M.E.Mabry & M.G.Simpson
Mexico USA
  var. purpusii Jepson
Mexico USA
  var. stenoloba I.M.Johnston
USA
Cryptantha utahensis (A.Gray) Greene
Mexico USA
     
Series Scoparieae *    
Cryptantha nevadensis A.Nelson & P.B.Kennedy
USA
Cryptantha scoparia A.Nelson 4.3
USA
     
Series Texanae (Johnston 1925)    
Cryptantha crassisepala (Torrey & A Gray) Greene
USA
   var. crassisepala
USA
   var. elachantha I.M.Johnston
  USA
Cryptantha fendleri (A.Gray) Greene 2B.2 *
Canada USA
Cryptantha kelseyana Greene
Canada USA
Cryptantha minima Rydberg
USA
Cryptantha texana (A.DC.) Greene
USA
     
Currently Unplaced to Series:    
Cryptantha excavata Brandegee 1B.1  WI
  USA
Cryptantha hooveri I.M.Johnston 1A
USA
Cryptantha recurvata Coville
USA
   
   
MARITIMAE CLADE [Simpson et al. 2017;
    Mabry and Simpson 2018; =Cryptantha s.s.2 of
    Hasenstab and Simpson 2012]
 
Series Maritimae (Johnston 1925)  
Cryptantha echinosepala J.F.Macbride
  Mexico  
Cryptantha maritima (Greene) Greene  SA
  Mexico USA
   var. cedrosensis (Greene) I. M. Johnston
Mexico
   var. maritima
Mexico USA
   var. pilosa I. M. Johnston  SA
Mexico USA
   var. vizcainensis Rebman & M.G.Simpson
Mexico  
Cryptantha pondii Greene
Mexico  
   
Series Muricatae (Johnston 1925)  
Cryptantha clokeyi I.M.Johnston  1B.2
USA
Cryptantha martirensis M. G. Simpson & Rebman
Mexico  
Cryptantha muricata (Hooker & Arnott) A.Nelson & J.F.Macbride
Mexico USA
   var. denticulata (Greene) I. M. Johnston
USA
   var. jonesii (A. Gray) I. M. Johnston
Mexico USA
   var. muricata
USA
   
NEW TAXA IN PROGRESS (names tentative):  
Cryptantha sp. nov.
Mexico USA
Cryptantha whippleae, sp. nov.
USA
TAXON/TAXÓN                                                                COUNTRY/PAÍS
SECTION EUCRYPTANTHA  
Series Capituliflorae (Johnston 1927)    
Cryptantha capituliflora (Clos) Reiche
Argentina Chile
Cryptantha longifolia (Philippi) Reiche
Chile
Cryptantha spathulata (Philippi) Reiche
Chile
     
Series Glomeratae (Johnston 1927)    
Cryptantha alfalfalis (Philippi) I.M.Johnston
Chile
Cryptantha aspera (Philippi) Grau
Chile
Cryptantha glomerata Lehmann ex G.Don
Chile
   subsp. glomerata
Chile
   subsp. quadrinuculata Grau
Chile
     
Series Glomeruliferae (Johnston 1927)    
Cryptantha glomerulifera (Philippi) I.M.Johnston
Argentina Chile
     
Series Haplostachyae (Johnston 1927)    
Cryptantha calycotricha I.M.Johnston
Chile
Cryptantha haplostachya (Philippi) I.M.Johnston
Chile
     
     
SECTION GEOCARYA    
Series Alyssoides (Johnston 1927)    
Cryptantha alyssoides (DC.) Reiche
Chile
   
Series Dimorphae (Johnston 1927)    
Cryptantha cynoglossoides (Philippi) I.M.Johnston [Grau 1981]
Argentina Chile
Cryptantha dimorpha (Philippi) Greene [Grau 1981]
Chile
Cryptantha involucrata (Philippi) Reiche
Chile
Cryptantha volckmannii (Philippi) I.M.Johnston
Chile
   
Series Dolichophyllae (Johnston 1927)    
Cryptantha dolichophylla (Philippi) Reiche
Chile
Cryptantha gayi I.M.Johnston Chile
Cryptantha macrocalyx (Philippi) Reiche
Chile
   
Series Lineares (Johnston 1927)  
Cryptantha aprica (Philippi) Reiche
Chile
Cryptantha linearis (Colla) Greene
Chile
   
Cryptantha chispae Grau
Chile
Cryptantha marticorenae Grau
Chile
   
   
SECTION "KRYNITZKIA" (not a monophyletic group)  
Series Barbigerae (not a monophyletic group)  
Cryptantha calycina (Philippi) Reiche
Chile
Cryptantha patagonica (Speg.) I.M.Johnston
Argentina
   
Series Gnaphalioides (Johnston 1927; Teillier 2009)  
Cryptantha gnaphalioides (A.DC.) Reiche
Chile
Cryptantha marioricardiana Teillier [tentative]
Chile
   
Series Phaceloides (Johnston 1927)  
Cryptantha dichita (Philippi) I.M.Johnston
Chile
Cryptantha hispida (Philippi) Reiche
Chile
Cryptantha phaceloides (Clos) Reiche
Chile
   
Series Texanae (Johnston 1925, 1930)  
Cryptantha mendocina I.M.Johnston
Argentina
   
Insertae Sedis  
Cryptantha latefissa R.L.Pérez-Mor.
Argentina
Cryptantha papillosa R.L.Pérez-Mor.
Argentina
   
   
GLOBULIFERA CLADE [Simpson et al. 2017;
    Mabry and Simpson 2018]
 
Cryptantha debilis (Philippi) Reiche
Bolivia Chile
Cryptantha diffusa (Philippi) I.M.Johnston
Argentina Bolivia Chile
Cryptantha globulifera (Clos) Reiche
Argentina Chile
Cryptantha peruviana I.M.Johnston
Bolivia Peru
Cryptantha werdermanniana I.M.Johnston
Chile
   
   
MARITIMAE CLADE [Simpson et al. 2017;
    Mabry and Simpson 2018; =Cryptantha s.s.2 of
    Hasenstab and Simpson 2012]
 
Series Maritimae (Johnston 1925)  
Cryptantha maritima (Greene) Greene  NA
Argentina
   var. pilosa I.M.Johnston  NA Argentina  
   
Cryptantha chaetocalyx (Philippi) I.M.Johnston
Chile
Cryptantha filaginea (Philippi) Reiche
Chile Peru
Cryptantha filiformis (Philippi) Reiche
Chile
Cryptantha granulosa (Ruiz & Pav.) I.M.Johnston
Peru
Cryptantha limensis (A.DC.) I.M.Johnston
Chile Peru
Cryptantha romanii I.M.Johnston
Chile
Cryptantha taltalensis I.M.Johnston
Chile
Cryptantha varians Brand   Peru
   
Cryptantha argentea I.M.Johnston
Chile
Cryptantha subamplexicaulis (Philippi) Reiche
Chile
   
UNKNOWN/UNCERTAIN  
Cryptantha unknown - South America  
   
   
California Native Plant Society (CNPS) Inventory Listings:
1A = California endemic, presumed extinct
1B = Rare, threatened, or endangered in California and elsewhere
2 = Rare, threatened, or endangered in California, but more common elsewhere
3 = review list (they need more information to list or not)
4 = Limited distribution (Watch List)
.1 = Seriously endangered in California (over 80% of occurrences threatened / high degree and immediacy of threat)
.2 = Fairly endangered in California (20-80% occurrences threatened)
.3 = Not very endangered in California (<20% of occurrences threatened or no current threats known)

Plant Taxa with High Affinity to Ultramafic Substrates in California (after Safford et al. 2005, Safford and Miller 2020):
BE/SI = Broad Endemic/Strong Indicator
SE = Strong Endemic
SI = Strong Indicator
WI = Weak Indicator
WI/IN = Weak Indicator/Indifferent



KEYS/CLAVES TO SOUTH AMERICAN CRYPTANTHA

CHROMOSOME NUMBERS OF AMSINCKIINAE, including Cryptantha




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