Cryptantha torreyana

Pronunciation/Pronunciación:	Cryp-tán-tha tor-re-yà-na
Etymology/Etimología:	after John Torrey (1796-1873)
Synonyms/Sinónimos:	Basionym: Krynitzkia torreyana A. Gray, Proc. Amer. Acad. Arts 20: 271. 1885 Homotypic Synonym(s): Cryptantha torreyana var. genuina I. M. Johnston, Contr. Gray Herb. 74: 78. 1925 Heterotypic Synonym(s): Krynitzkia torreyana A. Gray var. calycosa A. Gray, Proc. Amer. Acad. Arts 20: 271. 1885 Type: GH00097687 Cryptantha torreyana var. calycosa (A. Gray) Greene, Pittonia 1: 119. 1887 Cryptantha calycosa (A. Gray) Rydberg, Mem. New York Bot. Gard. 1: 331. 1900 Cryptantha affinis (A. Gray) Greene var. flexuosa A. Nelson, Bot. Gaz. 30: 195 1900 Cryptantha torreyi Rydberg, Mem. New York Bot. Gard. 1: 331 1900 Cryptantha pumila A. Heller, Muhlenbergia 2: 242 1906 Cryptantha flexuosa (A. Nelson) A. Nelson, Man. Bot. Rocky Mt. ed. 2: 416 1909 Cryptantha torreyana var. calistogae I. M. Johnston, Contr. Gray Herb. 74: 78. 1925 Holotype: Tracy 2094, 23 May 1904 (UC175784) Cryptantha torreyana var. genuina subvar. bracteata Brand, Pflanzenr. (Engler) 4, Fam. 252: 58. 1931 Cryptantha torreyana var. genuina subvar. calycosa Brand, Pflanzenr. (Engler) 4, Fam. 252: 58. 1931 Cryptantha torreyana var. genuina subvar. capitata Brand, Pflanzenr. (Engler) 4, Fam. 252: 58. 1931 Cryptantha eastwoodiae H. St. John, Fl. Southeast. Wash. & Idaho 342 (1937)
Type/Tipo:	Uncertain; listed as a possible type: E. L. Greene s.n., Jun 1887, USA: California, Alameda County, Berkeley (PH00016774)

Johnston 1925 Description, p. 78

Description in The Jepson eFlora

Phylogenetic Relationships:

Cladograms from: Top, Simpson et al. 2017:1411, cpDNA results; Bottom, Mabry and Simpson 2018:64, Astral results.
In all these results, Cryptantha torreyana (var. torreyana) is part of a well-supported clade, is sister to C. ambigua, these two sister to C. crinita and C. mariposae and these four sister to C. watsonii.
Note: There are undocumented reports that introgression or hybridization may occur between C. torreyana and C. ambigua.

Key to the two recognized varieties (after Kelley 2012 et al.)

  Calyx 1–1.5(2) mm, (2.5)3–4.5 mm in fruit, lobe margin sparse-strigose; nutlet 1.3–1.5 mm ................. C. t. var. pumila

  Calyx 2–2.5 mm, 4–6.5(8) mm in fruit, lobe margin dense-strigose; nutlet (1.6)1.7–2.2(2.5) mm ........ C. t. var. torreyana

Distribution of the species:

Kartesz, J.T., The Biota of North America Program (BONAP). 2012. North American Plant Atlas (http://www.bonap.org/MapSwitchboard.html).
Chapel Hill, N.C. [maps generated from Kartesz, J.T. 2010. Floristic Synthesis of North America, Version 1.0. Biota of North America Program (BONAP). (in press)].
COLOR KEY: Dark green=present in state; Light green=present in county, not rare; Yellow=present in county, rare. Accessed 12 Jan 2013.
Note: Cryptantha torreyana is the northern-most species in the genus.

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Cryptantha torreyana (A.Gray) Greene