Amsinckiinae Working Group

Nutlet image of Adelinia Nutlet image of Amsinckia Nutlet image of Amsinckiopsis Nutlet image of Andersonglossum Nutlet image of Cryptantha Nutlet image of Dasynotus Nutlet image of Eremocarya Nutlet image of Greeneocharis Nutlet image of Gruvelia Nutlet image of Harpagonella Nutlet image of Johnstonella Nutlet image of Oncaglossum Nutlet image of Oreocarya Nutlet image of Pectocarya Nutlet image of Plagiobothrys Nutlet image of Simpsonanthus Nutlet image of Sonnea
Adelinia Amsinckia Amsinckiopsis Andersonglossum Cryptantha Dasynotus Eremocarya Greeneocharis Gruvelia Harpagonella Johnstonella Oncaglossum Oreocarya Pectocarya Plagiobothrys Simpsonanthus Sonnea

Amsinckiinae Working Group

[Amsinckiinae HOME]

Matt Guilliams Kristen Haseenstab-Lehman Ron Kelley Makenzie Mabry Michael Simpson

Dr. Tim Böhnert

Nees Institute for
Biodiversity of Plants
University of Bonn,

Dr. C. Matt Guilliams

Department of Conservation
and Research
Santa Barbara Botanic Garden
Santa Barbara, California

Dr. Kristen Hasenstab-Lehman

Department of Conservation
and Research
Santa Barbara Botanic Garden
Santa Barbara, California

Dr. Ronald B. Kelley

Department of Chemistry
and Biochemistry
Eastern Oregon University
La Grande, Oregon

Dr. Makenzie E. Mabry

Florida Museum of Natural History
University of Florida
Gainesville, Florida

Dr. Pablo Moroni

Instituto de Botánica Darwinion CONICET - Academia Nacional de Ciencias Exactas
San Isidro, Buenos Aires

Dr. Jon P. Rebman

Department of Botany
San Diego Natural History Museum
San Diego, California

Dr. Michael G. Simpson

Department of Biology
San Diego State University
San Diego, California

      Amsinckiinae Working Group

  The goals of the AWG are to promote collaborative research into the phylogeny, taxonomy, and biogeography of the subtribe Amsinckiinae, Boraginaceae.


Balderis, E., C. M. Guilliams, K. E. Hasenstab-Lehman, M. E. Mabry, and M. G. Simpson. 2019. Amsinckiinae Working Group: Nutlet diversity of Amsinckiinae
      in western North America. Poster (pdf; png) presented at Botany 2019. Tucson, Arizona.
Chacón, J., F. Luebert, H. Hilger, S. Ovcinnikova, F. Selvi, L. Cecchi, C. M. Guilliams, K. Hasenstab-Lehman, K. K. Sutorý, M. G. Simpson, and M. Weigend. 2016.
      The borage family (Boraginaceae s.str.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some
      enigmatic genera. Taxon 3: 523-546.
Guilliams, C. M., B. A. Veno, M. G. Simpson, and R. B. Kelley. 2013. Pectocarya anisocarpa, a new species of Boraginaceae from western North America,
      with a revised dichotomous key and photographic atlas for the genus in the region. Aliso 31: 1-13.
Guilliams, C. M., T. Jang, and B. G. Baldwin. 2016. Molecular and morphological evidence for recognition of two species within Harpagonella (Amsinckiinae, Boraginaceae). PhytoKeys 70: 17-30.
Guilliams, M. C., K. Hasenstab-Lehman, M. Mabry, and M. G. Simpson. 2017. Memoirs of a frequent flier: Phylogenomics reveals 18 long-distance dispersals
     between North America and South America in the popcorn flowers (Amsinckiinae, Boraginaceae). American Journal of Botany 104: 1717-1728.
Guilliams, C. M., K. Hasenstab-Lehman, and B. G. Baldwin. 2020. Nomenclatural changes in western North American Amsinckiinae (Boraginaceae). Novon 28: 51-59.
Hargrove, L. and M. G. Simpson. 2003. Ultrastructure of heterocolpate pollen in Cryptantha (Boraginaceae). International Journal of Plant Sciences 164: 137-151.
Hasenstab-Lehman, K. E. and M. G. Simpson. 2012. Cat’s eyes and popcorn flowers: Phylogenetic systematics of the genus Cryptantha s.l.
     (Boraginaceae). Systematic Botany 37: 738–757.
Hasenstab-Lehman, K. E. 2017. Phylogenetics of the borage family: Delimiting Boraginales and assessing closest relatives. Aliso: A Journal of Systematic
      and Evolutionary Botany 31: 41-49.
Luebert, F., L. Cecchi, M. W. Frohlich, M. Gottschling, C. M. Guilliams, K. E. Hasenstab-Lehman, H. H. Hilger, J. S. Miller, M. Mittelbach, M. Nazaire, M. Nepi,
      D. Nocentini, D. Ober, R. G. Olmstead, F. Selvi, M. G. Simpson, K. Sutorý, B. Valdés, G. K. Walden, and M. Weigend. 2016. Familial classification of the
      Boraginales. Taxon 65: 502–522 .
Mabry, M. E., R. A. Dowdy, L. M. Simpson, J. P. Rebman, and M. G. Simpson. 2016. Taxonomy of the winged popcorn flower: Cryptantha pterocarya (Boraginaceae). Phytotaxa 253: 97-130.
Mabry, M. E. and M. G. Simpson. 2018. Evaluating the monophyly and biogeography of Cryptantha (Boraginaceae). Systematic Botany 43: 53-76.
Moroni, P., A. Martínez, and M. G. Simpson. 2021. Nomenclatural revision of Cryptantha (Boraginaceae s. str.) names linked to South American taxa. PhytoKeys 181:29-47.
Rebman, J. P. and M. G. Simpson. 2022. Three new species of Cryptantha (Boraginaceae) from the Southern Channel Islands of California. Madroño 69.
Ripma, L. A., M. G. Simpson, and K. Hasenstab-Lehman. 2014. Geneious! Simplified genome skimming methods for phylogenetic systematic studies:
      A case study in Oreocarya (Boraginaceae). Applications in Plant Sciences 2: 1400062.
Simpson, M. G. and K. E. Hasenstab. 2009. Cryptantha of Southern California. Crossosoma 35: 1-59.
Simpson, M. G. and J. P. Rebman. 2013. A new species of Cryptantha (Boraginaceae) from the Sierra de San Pedro Mártir, Baja California, Mexico. Madroño 60:35-45.
Simpson, M. G., J. P. Rebman, K. E. Hasenstab-Lehman, C. M. Guilliams, and P. O. McConnell. 2013. Cryptantha wigginsii (Boraginaceae): A presumed extinct species rediscovered.
      Madroño 60: 24-34.
Simpson, M. G., R. Dowdy, J. P. Rebman, R. B. Kelley, and L. M. Simpson. 2014. Recognition of two species in Eremocarya (Boraginaceae):
      Evidence from fornix bodies, nutlets, corolla size, and biogeography. Madroño 61: 259–275.
Simpson, M. G., K. Hasenstab-Lehman, and R. B. Kelley. 2014. Typification of the name Johnstonella (Boraginaceae). Taxon 63: 930-931.
Simpson, M. G., L. M. Simpson, and J. P. Rebman. 2016. A new, large-flowered variety of Eremocarya micrantha (Boraginaceae). Madroño 63: 39-54.
Simpson, M. G. and R. B. Kelley. 2017. Cryptantha nevadensis var. rigida elevated to species with a new name, C. juniperensis. Phytotaxa 295: 227-236.
Simpson, M. G., C. M. Guilliams, K. E. Hasenstab-Lehman, M. E. Mabry, and L. Ripma. 2017. Phylogeny of the popcorn flowers: Use of genome skimming to
    evaluate monophyly and interrelationships in subtribe Amsinckiinae (Boraginaceae). Taxon 66 :1406–1420.
Simpson, M. G., M. E. Mabry, and K. Hasenstab-Lehman. 2019. Transfer of four species of Cryptantha to the genus Johnstonella (Boraginaceae). Phytotaxa 425: 279–289.
Simpson, M. G., J. Stephens, and S. Yang. 2020. Noteworthy Collections: Johnstonella angelica (I.M.Johnston) Hasenstab & M.G.Simpson. Madroño 67: 5–8.
Simpson, M. G. and J. P. Rebman. 2021. Research in Boraginaceae: A new variety of Cryptantha maritima, Cryptantha pondii resurrected, and Johnstonella echinosepala transferred back to Cryptantha.
Phytotaxa 509:185–210.
Simpson, M.G. & Rebman, J.P. 2021. A new species of Cryptantha restricted to dunes in northwestern Baja California, Mexico. Madroño 68: 126-136.
Simpson, M. G., K. E. Hasenstab-Lehman, M. E. Mabry, and R. B. Kelley. 2022. Jepson eFlora: Cryptantha. Jepson Flora Project (eds.) 2022.

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