Nutlet image of Adelinia Nutlet image of Amsinckia Nutlet image of Amsinckiopsis Nutlet image of Andersonglossum Nutlet image of Cryptantha Nutlet image of Dasynotus Nutlet image of Eremocarya Nutlet image of Greeneocharis Nutlet image of Gruvelia Nutlet image of Harpagonella Nutlet image of Johnstonella Nutlet image of Oncaglossum Nutlet image of Oreocarya Nutlet image of Pectocarya Nutlet image of Plagiobothrys Nutlet image of Simpsonanthus Nutlet image of Sonnea
Adelinia Amsinckia Amsinckiopsis Andersonglossum Cryptantha Dasynotus Eremocarya Greeneocharis Gruvelia Harpagonella Johnstonella Oncaglossum Oreocarya Pectocarya Plagiobothrys Simpsonanthus Sonnea

Systematics of Amsinckiinae (Boraginaceae): The Popcorn Flowers

[Amsinckiinae HOME]

Harpagonella A.Gray

Proc. Amer. Acad. Arts 11: 88 (1876)

diminuitive of harpago, a small grappling hook, from the calyx prickles

Plagiobothrys Fisch. & C.A.Mey., Index Seminum [St.Petersburg (Petropolitanus)] 2: 46 (printed as 21) (1836)
   Veno (1979:60) proposed transferring Harpagonella palmeri to Pectocarya, as P. palmeri (A.Gray) Veno,
    but this was not validly published.

Type Species/EspeciesTipo:
Harpagonella palmeri A.Gray, Proc. Amer. Acad. Arts 11: 88 (1876)

Distribution of Harpagonella:
Harpagonella distribution from POWO Harpagonella distribution from GBIF.org 
POWO. Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet. Green=native.
MIDDLE: GBIF. Global Biodiversity Information Facility.
RIGHT: SEINet. Click to see current herbarium vouchers. Note: Specimens in Arizona are likely all H. arizonica.

2 species and 2 minimum ranked taxon
North American only (USA and Mexico)

California Native Plant Society (CNPS) Inventory Listings:
1A = Plants presumed extirpated in California and either rare or extinct elsewhere
1B = Plants rare, threatened, or endangered in California and elsewhere
2A = Plants presumed extirpated in California but common elsewhere
2B = Plants rare, threatened, or endangered in California but common elsewhere
3 = Plants about which more information is needed, a review list
4 = Plants of limited distribution, a watch list
.1 = Seriously threatened in California — Over 80% of occurrences threatened / high degree and immediacy of threat
.2 = Moderately threatened in California — 20-80% of occurrences threatened / moderate degree and immediacy of threat
.3 = Not very threatened in California — Less than 20% of occurrences threatened / low degree and immediacy of threat or no current threats known

KEY TO TAXA (after Guilliams et al. 2016)

1. Fruits 6-9 mm long, appendages (average) 3.3-5 mm long; Arizona and northern Sonora (rarely Baja California), Mexico ......... H. arizonica
1' Fruits 3-6 mm long, appendages (average) 1.5-3.2 mm long; southwestern California and Baja California peninsula, Mexico .....
H. palmeri



Gray, A. 1876. V. Miscellaneous botanical contributions. Proceedings of the American Academy of Arts and Sciences 11:71-104.
Guilliams, C. M., T. Jang, and B. G. Baldwin. 2016. Molecular and morphological evidence for recognition of two species within Harpagonella (Amsinckiinae, Boraginaceae).
      PhytoKeys 70: 17-30.
Johnston, I.M. 1924. Studies in the Boraginaceae II. A synopsis of the American native and immigrant Borages of the subfamily Boraginoideae. Contr. Gray Herb. 70: 3-61.
Veno, B. A. 1979. A revision of the genus Pectocarya (Boraginaceae) including reduction to synonymy of the genus Harpagonella (Boraginaceae). Doctoral Dissertation,
     University of California, Los Angeles.

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